Question

What is the evolutionary advantage of closure of the septum between the two ventricles to create a double circulatory system?

Short answer

The evolutionary advantage of a closed septum resulting in a double circulatory system includes more efficient oxygenation of blood, higher metabolic rates, improved nutrient delivery, and support for high levels of activity and endothermy.

Step-by-step solution

Understanding the Septum

The septum is a dividing wall that separates the right and left ventricles of the heart. In reptiles and amphibians, the septum is not complete, allowing for the mixing of oxygenated and deoxygenated blood. In mammals and birds, however, the septum is complete, which prevents this mixing and leads to a more efficient double circulatory system.

Advantages of Complete Septum

The closure of the septum offers several evolutionary advantages. It provides separation of the pulmonary (lung) and systemic (body) circulation, allowing for higher blood pressure in the systemic circuit. This separation results in more efficient oxygenation of the blood, higher metabolic rates, and improved delivery of oxygen and nutrients to tissues, which is especially beneficial for warm-blooded, high-energy-requiring organisms like mammals and birds.

Result of Double Circulation

The double circulatory system with a closed septum enables a faster and more efficient transport of oxygen and carbon dioxide between the lungs and the rest of the body. This efficient system supports higher levels of activity and endothermy (regulation of body temperature), providing a significant evolutionary advantage for survival and diversification in various environments.

Key concepts

Septum Closure in the Heart

The septum within the heart is key to understanding the benefits of a double circulatory system. This wall's main function is to partition the heart into two separate sides. The evolutionary transition that led to a closed septum in mammals and birds marks a significant upgrade in circulatory efficiency.

In creatures with an incomplete septum—think amphibians and reptiles—oxygen-rich blood can mix with oxygen-poor blood, resulting in a less efficient system. With the septum's closure, each side of the heart handles distinct types of blood: one oxygenated, the other not. By preventing the mix, the animal's body ensures that tissues receive the most oxygenated blood possible.

Efficiency and High Energy Demands

The separation brought about by a complete septum satisfies the high energy demands of warm-blooded animals. Since they maintain a constant body temperature, the energy requirements are substantial, making the efficient use of oxygen imperative.

Pulmonary and Systemic Circulation

Understanding the way blood circulates through the body is critical. In a double circulatory system, blood travels through the heart twice in one complete circuit of the body. There are two distinct loops: pulmonary circulation and systemic circulation.

Pulmonary Circulation

This loop sends deoxygenated blood from the right side of the heart to the lungs to pick up oxygen. The blood then returns to the heart, distinctly richer in oxygen.

Systemic Circulation

Now oxygenated, the blood ventures from the left side of the heart to nourish all body tissues. It delivers oxygen and nutrients, collects waste, and then, lacking oxygen again, heads back to the heart.

By having separate pathways for oxygenated and deoxygenated blood, the body maintains higher energy levels and efficiency. This is particularly useful for animals with high metabolic rates.

Endothermy in Mammals and Birds

Endothermy, or warm-bloodedness, is the ability to regulate one's internal temperature through metabolic processes. Mammals and birds, both endotherms, are in a constant battle to keep their body temperatures stable, irrespective of external conditions. This heat production requires a significant amount of energy.

High Metabolic Rates

These animals have high metabolic rates, which require a relentless supply of oxygen to fuel the countless reactions that produce heat. The double circulatory system, empowered by a fully enclosed septum, efficiently provides oxygen-rich blood to tissues and organs.

Sustaining a steady body temperature is vital for survival. It aids in the efficiency of enzymatic reactions, allowing mammals and birds to be active across a wide range of temperatures. It's a clear evolutionary advantage that supports their diverse lifestyles and ecological roles.